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To prevent endocytosis, metabolic inhibitors (10 mM NaN 3 , 20 mM 2-deoxyglucose) 9 were added 20 min before labeling completion and maintained throughout the experiment when indicated

Merck Millipore, Darmstadt, Germany) tubes and centrifuged at 75,000 g for 40 min at 4 C

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