The GCH1/BH4/DHFR axis operates independently of GPX4: guanosine triphosphate cyclohydrolase 1 (GCH1) catalyzes the rate-limiting step in the de novo synthesis of tetrahydrobiopterin (BH4) from GTP, producing BH4 as a potent radical-trapping antioxidant (RTA) that directly neutralizes lipid peroxyl radicals (LOO) and prevents membrane lipid peroxidation ( 2 ) a lipophilic radical-trapping antioxidant that detoxifies lipid peroxides where GPX4 access is limited
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In 1919 the painting was bought via the dealers Colnaghi's for 60,000 by Henry Lascelles, 6th Earl of Harewood (then Viscount Lascelles), giving "the measure of Rothschild's expertise and of Brownlow's generosity"
